All cats descend from the Felidae family, sharing similar musculature, cardiovascular systems, skeletal frames, and behaviour. Both the cheetah and cougar differ physically from fellow big cats, and to a greater extent, other small cats. As obligate carnivores, big cats are considered apex predators, topping their food chain without natural predators of their own. Native ranges include the Americas, Africa, and Asia; the range of the leopard and tiger also extends into Europe, specifically in Russia.
It is estimated that the ancestors of most big cats split away from the Felinae about 6.37 million years ago. The Felinae, on the other hand, comprises mostly small to medium-sized cats, including the domestic cats, but also some larger cats such as the cougar and cheetah.
A 2010 study published in Molecular Phylogenetics and Evolution has given insight into the exact evolutionary relationships of the big cats. The study reveals that the snow leopard and the tiger are sister species, while the lion, leopard, and jaguar are more closely related to each other. The tiger and snow leopard diverged from the ancestral big cats approximately 3.9 Ma. The tiger then evolved into a unique species towards the end of the Pliocene epoch, approximately 3.2 Ma. The ancestor of the lion, leopard, and jaguar split from other big cats from 4.3–3.8 Ma. Between 3.6–2.5 Ma the jaguar diverged from the ancestor of lions and leopards. Lions and leopards split from one another approximately 2 Ma. The earliest big cat fossil, Panthera blytheae, dating to 4.1−5.95 MA, was discovered in southwest Tibet.

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