The Seattle Pilots drafted Coluccio in the 17th round of the 1969 MLB Draft out of Centralia High School. Coluccio chose to sign with the Pilots for $500 monthly (equivalent to $3,695 in 2021) instead of operating a paver for his father's company, Civic Sand & Gravel. He began his career with the Billings Mustangs.
Coluccio took a hiatus in 1971 to get married, during which time his contract was optioned to the Philadelphia Phillies. He finished the 1971 season with the Reading Phillies. He returned to the Milwaukee Brewers system the following season.
Coluccio was invited to Major League spring training by the Brewers in 1973 following an injury to outfielder Dave May and broke camp with the team. The first MLB game which Coluccio attended was Opening Day at Milwaukee County Stadium in 1973. Coluccio became popular with Milwaukee's Italian-American population and was nicknamed the "Macaroni Pony" by Milwaukee Brewers broadcasters Bob Uecker and Merle Harmon.

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