Melendez was nicknamed after the El Niño storm, it was given to him by a Japanese broadcaster describing his fight-style and the manner in which he stormed through the Shooto Lightweight division.
Melendez is from Santa Ana, California, of Mexican descent. He wrestled at Santa Ana High School. Melendez always enjoyed combat sports and competition, but did not start training in MMA until after college when he was introduced to the sport by his good friend and MMA fighter Jake Shields. Melendez attended San Francisco State University and was studying in liberal arts, he anticipated becoming a teacher but did not finish his college education and get a degree, on account of training for MMA and working outside of college. He has stated that someday he plans on finishing his college education. Before becoming a professional fighter, he worked as a waiter and he also had a job at a Fairtex gear shop, which would later be one of his sponsors.
Melendez made his debut in the World Extreme Cagefighting and started off his career dominating his opponents, winning his first three career fights all by knockout due to punches before receiving an opportunity to fight for the first WEC Lightweight Championship against fan-favorite Olaf Alfonso at WEC 10. Melendez won the bout by TKO due to punches and became the first WEC Lightweight Champion. Two fights later, Melendez, still undefeated, made his debut in Japan's Shooto against Hiroyuki Takaya, which Melendez won by unanimous decision. He followed this up with two more TKO wins in Shooto to bring his record to 8-0 before making his Strikeforce debut against Harris Sarmiento, which Melendez won after his opponent tapped out due to punches received.

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